These species belong to Copris, a genus of dung beetles in the family Scarabaeidae.[1][2]

Copris species

  • Copris acutidens Motschulsky, 1860
  • Copris afgoi Cambefort, 1992
  • Copris agnus Sharp, 1875
  • Copris algol Nguyen-phung, 1989
  • Copris amabilis Kolbe, 1914
  • Copris amyntor Klug, 1855
  • Copris anceus (Olivier, 1789)
  • Copris andrewesi Waterhouse, 1891
  • Copris angusticornis Arrow, 1933
  • Copris angustus Nguyen-phung & Cambefort, 1986
  • Copris anomiopseoides Boucomont, 1924
  • Copris antares Ferreira, 1958
  • Copris arcturus Gillet, 1907
  • Copris ares Zelenka, 1993
  • Copris arimotoi Ochi & Araya, 1992
  • Copris arizonensis Schaeffer, 1906
  • Copris armatus Harold, 1869
  • Copris armeniacus Faldermann, 1835
  • Copris armiger Gillet, 1910
  • Copris arrowi Felsche, 1910
  • Copris aspericollis Gillet, 1910
  • Copris atropolitus Gillet, 1910
  • Copris bacchus Thunberg, 1787
  • Copris barclayi Ochi, Kon & Bai, 2009
  • Copris basilewskyi Ferreira, 1962
  • Copris basipunctatus Balthasar, 1942
  • Copris bellator Chevrolat, 1844
  • Copris bengalensis Gillet, 1911
  • Copris bidens Kolbe, 1893
  • Copris binodis
  • Copris bootes Klug, 1855
  • Copris boucardi Harold, 1869
  • Copris bovinus Gillet, 1908
  • Copris brachypterus Nomura, 1964
  • Copris brittoni Ferreira, 1962
  • Copris caelatus (Fabricius, 1794)
  • Copris caliginosus Kohlmann, Cano & Delgado, 2003
  • Copris cambeforti Nguyen-phung, 1988
  • Copris cambodiensis Ochi, Kon & Kawahara, 2008
  • Copris camerunus Felsche, 1904
  • Copris caobangensis Bui, Dumack & Bonkowski, 2018 [3]
  • Copris capensis Waterhouse, 1891
  • Copris cariniceps Felsche, 1910
  • Copris carinicus Gillet, 1910
  • Copris carmelita Fabricius, 1801
  • Copris cassius Péringuey, 1901
  • Copris celebensis Lansberge, 1886
  • Copris cheni Ochi, Kon & Bai, 2007
  • Copris chimalapensis Mora-Aguilar & Delgado, 2015
  • Copris ciliatus
  • Copris colmanti Gillet, 1908
  • Copris complexus Nguyen-phung, 1989
  • Copris compressipennis Gillet, 1910
  • Copris confucius Harold, 1877
  • Copris coriarius Gillet, 1907
  • Copris cornifrons Boheman, 1860
  • Copris corniger Sahlberg, 1823
  • Copris corpulentus Gillet, 1910
  • Copris costaricensis Gahan, 1894
  • Copris costariensis Matthews
  • Copris crassus Deschodt & Davis, 2015
  • Copris cribratus Gillet, 1927
  • Copris cribricollis Gillet, 1910
  • Copris curlettii Moretto, 2011
  • Copris davidis (Deyrolle, 1878)
  • Copris davisi Nguyen-phung & Cambefort, 1986
  • Copris davisoni Waterhouse, 1891
  • Copris decellei Cambefort, 1992
  • Copris delicatus Arrow, 1931
  • Copris denticulatus Nguyen-phung, 1988
  • Copris diversus Waterhouse, 1891
  • Copris draco Arrow, 1906
  • Copris dracunculus Fereira, 1959
  • Copris druidum Heer, 1862
  • Copris dudleyi Cambefort, 1992
  • Copris eburneus Cambefort, 1992
  • Copris elpheneroides Felsche, 1910
  • Copris elphenor Klug, 1855
  • Copris erratus Lansberge, 1886
  • Copris evanidus Klug, 1855
  • Copris excisus Waterhouse, 1891
  • Copris fallaciosus Gillet, 1907
  • Copris fallax Felsche, 1910
  • Copris felschei Reitter, 1892
  • Copris fidius (Olivier, 1789)
  • Copris frankenbergerianus Balthasar, 1958
  • Copris freyi Ferreira, 1961
  • Copris fricator (Fabricius, 1787)
  • Copris fukiensis Balthasar, 1952
  • Copris furciceps Felsche, 1910
  • Copris garambae Cambefort, 1992
  • Copris gazellarum Gillet, 1918
  • Copris gibbulus Lansberge, 1886
  • Copris gilleti Kolbe, 1907
  • Copris gladiator Arrow, 1933
  • Copris gopheri Hubbard, 1894
  • Copris gracilis Waterhouse, 1891
  • Copris graueri Kolbe, 1914
  • Copris halffteri Matthews, 1959
  • Copris harrisi Waterhouse, 1891
  • Copris hartli Frey, 1975
  • Copris hatayamai Ochi & Araya, 1992
  • Copris hispanus (Linnaeus, 1764)
  • Copris howdeni Matthews & Halffter, 1959
  • Copris humilis Gillet, 1908
  • Copris hybridus Gillet, 1910
  • Copris igualensis Warner, 1990
  • Copris illotus Balthasar, 1942
  • Copris imitans Felsche, 1910
  • Copris inaequabilis Zhang, 1997
  • Copris incertus Say, 1835
  • Copris inemarginatus Blatchley, 1918
  • Copris inhalatus Quedenfeldt, 1884
  • Copris insidiosus Péringuey, 1901
  • Copris integer Reiche, 1847
  • Copris interioris Kolbe, 1897
  • Copris iris Sharp, 1875
  • Copris jacchoides Nguyen-phung & Cambefort, 1987
  • Copris jacchus (Fabricius, 1775)
  • Copris jahi Nguyen-phung & Cambefort, 1986
  • Copris johkii Ochi & Kon, 2014
  • Copris juanmorai Mora-Aguilar & Delgado, 2015
  • Copris jucundus Gillet, 1932
  • Copris kachinensis Ochi, Kon & Kawahara, 2008
  • Copris kartlinus Kabakov, 1988
  • Copris kasagii Ochi & Kon, 1996
  • Copris kasanka Josso & Prévost, 2016
  • Copris katangae Cambefort & Nguyen-phung, 1996
  • Copris keralensis Gill, 1986
  • Copris kiuchii Masumoto, 1989
  • Copris klapperichi Balthasar, 1942
  • Copris klugi Harold, 1869
  • Copris kusakabei Ochi, Kon & Kawahara, 2005
  • Copris laeviceps Harold, 1862
  • Copris laevigatus Gillet, 1927
  • Copris laiiformis Nguyen-phung & Cambefort, 1987
  • Copris laioides Boucomont, 1932
  • Copris laius Harold, 1868
  • Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
  • Copris leakeyorum Paulian, 1976
  • Copris lecontei Matthews, 1962
  • Copris leruei Josso & Prévost, 2017
  • Copris lugubris Boheman, 1858
  • Copris lunaris (Linnaeus, 1758)
  • Copris macacus Lansberge, 1886
  • Copris macclevei Warner, 1990
  • Copris macer Péringuey, 1901
  • Copris maesi Ratcliffe, 1998
  • Copris magicus Harold, 1881
  • Copris manni Ochi, Kon & Bai, 2009
  • Copris marcus Gillet, 1921
  • Copris martinae Nguyen-phung, 1988
  • Copris martinezi Matthews and Halffter, 1968
  • Copris matthewsi Delgado & Kohlmann, 2001
  • Copris medogensis Zhang, 1988
  • Copris megaceratoides Waterhouse, 1891
  • Copris megasoma Matthews & Halffter, 1959
  • Copris mesacanthus Harold, 1878
  • Copris mexicanus Matthews & Halffter, 1959
  • Copris minutus (Drury, 1773)
  • Copris misellus Péringuey, 1901
  • Copris moechus LeConte, 1854
  • Copris moffartsi Gillet, 1907
  • Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
  • Copris montivagus Gillet, 1908
  • Copris morphaeus Gillet, 1932
  • Copris mourgliai Zelenka, 1993
  • Copris neglectus Moxey, 1963
  • Copris neotridens Génier & Prévost, 2017
  • Copris nepos Gillet, 1908
  • Copris nevinsoni Waterhouse, 1891
  • Copris novaki Zelenka, 1992
  • Copris nubilosus Kohlmann & Cano & Delgado, 2003
  • Copris numa Lansberge, 1886
  • Copris obenbergeri Balthasar, 1933
  • Copris obesus Boheman, 1857
  • Copris ochus (Motschulsky, 1860)
  • Copris orion Klug, 1835
  • Copris orphanus Guérin-Méneville, 1847
  • Copris parapecuarius Ochi, Kon & Kawahara, 2008
  • Copris pauliani Nguyen-phung & Cambefort, 1986
  • Copris pecuarius Lewis, 1884
  • Copris pedarioides Lansberge, 1886
  • Copris phungae Cambefort, 1992
  • Copris phylax Gillet, 1908
  • Copris poggii Ochi & Kon, 2005
  • Copris potanini Semenov, 1891
  • Copris prevosti Josso, 2011
  • Copris pristinus Pierce, 1946
  • Copris pseudobootes Ferreira, 1962
  • Copris pseudochus Ochi & Kon, 2004
  • Copris pseudomoffartsi Moretto, 2011
  • Copris pseudosinicus Balthasar, 1958
  • Copris pueli Mollandin De Boissy, 1905
  • Copris punctatus Gillet, 1910
  • Copris puncticollis Boheman, 1857
  • Copris punctipennis Boucomont, 1914
  • Copris punctulatus Wiedemann, 1823
  • Copris punjabensis Gillet, 1921
  • Copris quasilaevigatus Ochi, Kon & Bai, 2007
  • Copris ramosiceps Gillet, 1921
  • Copris rebouchei Harold, 1869
  • Copris remotus Leconte, 1866
  • Copris renwarti Nguyen-phung & Cambefort, 1986
  • Copris repertus Walker, 1858
  • Copris ribbei Lansberge, 1886
  • Copris riekoae Ochi & Kon, 2003
  • Copris ritsemae Harold, 1875
  • Copris rugosus Gillet, 1908
  • Copris ruricola Balthasar, 1933
  • Copris sabinus Gillet, 1910
  • Copris sacontala Redtenbacher, 1848
  • Copris sallei Harold, 1869
  • Copris sarpedon Harold, 1868
  • Copris saundersi Harold, 1869
  • Copris schoolmeestersi Ochi & Kon, 2010
  • Copris scorpio Zelenka, 1993
  • Copris serius Nguyen-phung, 1987
  • Copris sexdentatus Thunberg, 1818
  • Copris siamensis Gillet, 1910
  • Copris siangensis Biswas, 1980
  • Copris sierrensis Matthews, 1961
  • Copris signatus Walker, 1858
  • Copris simonettai Cambefort, 1992
  • Copris simulator Nguyen-phung & Cambefort, 1986
  • Copris singularis Nguyen-phung & Cambefort, 1986
  • Copris sinicus Hope, 1842
  • Copris sinon (Olivier, 1789)
  • Copris sodalis Walker, 1858
  • Copris sonensis Bui, Dumack & Bonkowski, 2018 [3]
  • Copris sorex Balthasar, 1942
  • Copris sphaeropterus Harold, 1877
  • Copris spinator Harold, 1881
  • Copris subdolus Balthasar, 1958
  • Copris subpunctatus Gillet, 1910
  • Copris subsidens Péringuey, 1901
  • Copris subterraneus Heer, 1862
  • Copris sumatrensis Gillet, 1921
  • Copris surdus Arrow, 1931
  • Copris szechouanicus Balthasar, 1958
  • Copris tetraodon Gillet, 1910
  • Copris tridentatus Solis & Kohlmann, 2003
  • Copris tripartitus Waterhouse, 1875
  • Copris truncatus Felsche, 1901
  • Copris tsukamotoi Ochi, Kon & Kawahara, 2008
  • Copris typhoeus Gerstaecker, 1884
  • Copris uenoi Hanboonsong & Masumoto & Ochi, 2003
  • Copris umbilicatus Abeille De Perrin, 1901
  • Copris urus Boheman, 1857
  • Copris usambaricus Gillet, 1908
  • Copris vankhaii Nguyen-phung, 1988
  • Copris victorini Boheman, 1857
  • Copris vietnamicus Kabakov, 1994
  • Copris vilhenai Ferreira, 1962
  • Copris vrydaghi Ferreira, 1962
  • Copris warneri Mccleve & Kohlmann, 2005
  • Copris wiesei Kolbe, 1914
  • Copris yangi Ochi, Kon & Bai, 2007
  • Copris youngai Balthasar, 1967
  • Copris zhangi Ochi, Kon & Bai, 2009

References

  1. "Copris". GBIF. Retrieved 2019-01-05.
  2. "Copris Genus Information". BugGuide.net. Retrieved 2019-01-05.
  3. 1 2 Bui, Van Bac; Dumack, Kenneth; Bonkowski, Michael (2018). "Two new species and one new record for the genus Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Vietnam with a key to Vietnamese species". European Journal of Entomology. 15. doi:10.14411/eje.2018.016.
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